{# load django_bootstrap5#%} CCDB server
Prof. Itay Mayrose Lab - Plant Evolution, bioinformatics, & comparative genomics


   CCDB is a community resource. If you are aware of any missing data or if any of the counts are erroneous please submit it to the forms submit new counts, or report erroneous counts.


CCDBAngiospermsApiaceaeHeracleumHeracleum sphondylium

110 chromosome counts in Heracleum sphondylium:


Name Accepted Name Gametophytic(n) Sporophytic(2n) Data Source reference
  Heracleum alpinum L. Heracleum sphondylium subsp. alpinum (L.) Bonnier & Layens   22+0-1B IPCN online Tonascia, N. 1992. Biosystematische Untersuchungen an Heracleum sphondylium s.l. in der Schweiz. Ber. Geobot. Inst. ETH Stiftung Rübel 58: 101–120.
  Heracleum dissectum Ledeb. Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IAPT/IOPB IAPT/IOPB Chromosome Data 6
  Heracleum dissectum Ledeb. Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IPCN online Krogulevich, R. E. 1978. Kariologicheskij analiz vidov flory Vostochnogo Sajana. V Flora Pribajkal'ja. 19–48. Nauka, Novosibirsk.
  Heracleum dissectum Ledeb. Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IPCN online Rostovtseva, T. S. 1979. Chromosome numbers of some species of the family Apiaceae Lindl. II. Bot. Zhurn. 64 (2): 227–232.
  Heracleum dissectum Ledeb. Heracleum sphondylium subsp. montanum (Schleich.) Briq. 11   IPCN online Rostovtseva, T. S. 1976. Chromosome numbers of some species of the family Apiaceae in South Siberia. Bot. Žhurn. (Moscow & Leningrad) 61(1): 93–99.
  Heracleum dissectum Ledeb. Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IPCN online Krogulevich, R. E. 1978. Karyological analysis of the species of the flora of eastern Sayana. Pp. 19-48 in L. I. Malyshev & G. A. Peshlcova (eds.) Flora of the Prebaikal. Novosibirsk.
  Heracleum dissectum Ledeb. Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IAPT/IOPB IAPT/IOPB Chromosome Data 18
  Heracleum granatense Boiss. Heracleum sphondylium subsp. granatense (Boiss.) Briq.   22 IPCN67-71 CAUWET, A. - M. 1968. Contriburion a l'etude caryologique des Ombelliferes de la partie orientale des Pyrenees. Naturalia Monspel. Ser. Bot. 19: 5-27.
  Heracleum montanum Schleich. ex Gaudin Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IPCN online Tonascia, N. 1992. Biosystematische Untersuchungen an Heracleum sphondylium s.l. in der Schweiz. Ber. Geobot. Inst. ETH Stiftung Rübel 58: 101–120.
  Heracleum orsinii Guss. Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer   22 CHROBASE BALTISBERGER M. - 1991 (1990). Numeri cromosomici per la flora italiana: 1208-1230.. Inform. Bot. Ital. , 22(3): 216-226.
  Heracleum orsinii Guss. Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer   22 IPCN online Baltisberger, M. 1990. Numeri cromosomici per la flora Italiana: 1208--1230. Inform. Bot. Ital. 22: 216–226.
  Heracleum palmatum Baumg. Heracleum sphondylium subsp. transsilvanicum (Schur) Brummitt 11   IPCN66 Bell & Constance 1966
  Heracleum palmatum Baumg. Heracleum sphondylium subsp. transsilvanicum (Schur) Brummitt   22 Fedorov, 1974 Bell, Constance 1966
  Heracleum pollinianum Bertol. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 CHROBASE TONASCIA N. - 1992. Biosystematic investigations on Heracleum sphondylium s.l. in Switzerland.. Ber. Geobot. Inst. E.T.H., Stiftung Rubel , 58: 101-120.
  Heracleum pollinianum Bertol. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 IPCN online Tonascia, N. 1992. Biosystematische Untersuchungen an Heracleum sphondylium s.l. in der Schweiz. Ber. Geobot. Inst. ETH Stiftung Rübel 58: 101–120.
  Heracleum pyrenaicum Lam. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 IPCN online Baltisberger, M. 1999. IOPB chromosome data 15. Newslett. Int. Organ. Pl. Biosyst. (Pruhonice) 31: 11–12.
  Heracleum pyrenaicum Lam. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 IPCN67-71 CAUWET, A. - M. 1968. Contriburion a l'etude caryologique des Ombelliferes de la partie orientale des Pyrenees. Naturalia Monspel. Ser. Bot. 19: 5-27.
!   Heracleum setosum Lapeyr. Heracleum sphondylium subsp. montanum (Schleich.) Briq. 11   IPCN67-71 CAUWET, A. - M. 1968. Contriburion a l'etude caryologique des Ombelliferes de la partie orientale des Pyrenees. Naturalia Monspel. Ser. Bot. 19: 5-27.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 Poland Database Jankun A., Ma?ecka J., Izmai?ow R., Wcis?o H., Czapik R., Musia? K., et al. 1996 Further studies in chromosome numbers of Polish Angiosperms. Part XXV. Acta Biol. Cracov. Ser. Bot. 38, 9-27.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online Nanushyan, E. R., N. L. Bolsheva, N. S. Badaev & V. N. Tikhomirov. 1985. Differential staining of Heracleum sibiricum L. chromosomes with the help of C-method. Biol. Nauki (Moscow) 5(257): 72–75.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online Rostovtseva, T. S. 1982. The chromosome numbers of some species of the family Apiaceae. III. Bot. Zhurn. SSSR 67 (2): 206–210. (In Russian).
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online Parfenov, V. I. & S. A. Dmitrieva. 1987. Kariologicheskaja kharakteristika predstavitelej flory sosudistykh rastenij Berezinskogo Biosfernogo Zapovednika. II. Zapov. Belorussii Issl. 11: 62–69.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online Nikolov, N. A. 1991. Chromosome numbers of Bulgarian angiosperms from North Pirin Mountain: Reserve "Bajuvi Dupki--D|3zind|3zirica". Fitologija 41: 70–75.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online Dmitrieva, S. A., V. I. Parfenov & I. V. SCHVEC. 1977. The karyological characterization of some species of the Belorussian flora. Vesci Akad. Navuk Belarusk SSR Ser. Bi jal. Sel'skagasp. Navuk. 19: 82–91.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online GAGNIDZE, R. J. & P. B. CHKHEIDZE. 1974. The chromosome numbers of some species of Caucasian flora. Soobshch. Akad. Nauk Gruzinsk. SSR 75: 681–684.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk. 11   IPCN66 Bell & Constance 1966
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN67-71 KUZMANOV, B., & s. KOZUHAROV. 1969. Chromosome numbers of flowering plants in Bulgaria. 2. Izv. Bot. Inst. (Sofia) 19: 109-115.
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 Fedorov, 1974 Bell, Constance 1966
  Heracleum sibiricum L. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IAPT/IOPB IAPT/IOPB Chromosome Data 27
!   Heracleum sphondylium Heracleum sphondylium L. 11   IPCN online Wetschnig, W. & G. H. Leute. 1991. Chromosomenzahlen Kärntner Gefässflanzen (Teil 2, Doldenblütler---Apiaceae = Umbelliferae). Linzer Biol. Beitr. 23: 457–481.
!   Heracleum sphondylium Heracleum sphondylium L.   22+1S IPCN online Dempsey, R. E., R. J. Gornall & J. P. Bailey. 1994. Contributions to a cytological catalogue of the British and Irish flora, 4. Watsonia 20: 63–66.
!   Heracleum sphondylium Heracleum sphondylium L. 11 22 IPCN online Weimarck, G., F. Stewart & J. Grace. 1979. Morphometric and chromatographic variation and male meiosis in the hybrid Heracleum mantegazzianum X H. sphondylium (Apiaceae) and its parents. Hereditas 91: 117–127.
!   Heracleum sphondylium Heracleum sphondylium L.   22 IPCN online Murin, A. 1974. In Index of chromosome numbers of Slovakian flora. Part 4. Acta Fac. Rerum Nat. Univ. Comenianae, Bot. 23: 1–23.
!   Heracleum sphondylium Heracleum sphondylium L. n = 11   Austria_checklist Wetschnig, W. & Leute, G. H. 1991. Chromosomenzahlen K?rntner Gef??pflanzen. Teil 2, Doldenbl?tler - Apiaceae = Umbelliferae. Linzer Biol.Beitr. 23[2], 457-481.°
!   Heracleum sphondylium Heracleum sphondylium L. n = 11 II + 1 B I   Austria_checklist -- 1978. B chromosomes in Heracleum sphondylium s. lat. in Europe. Cauwet-Marc, A. M. and Carbonnier, J. Les Ombellif?res. Contributions pluridisciplinaires ? la systematique. pp. 293-303. Actes du 2e symposium international sur les Ombelliferes. Centre
!   Heracleum sphondylium Heracleum sphondylium L.   2n = 22 + 2 B* Austria_checklist -- 1978. B chromosomes in Heracleum sphondylium s. lat. in Europe. Cauwet-Marc, A. M. and Carbonnier, J. Les Ombellif?res. Contributions pluridisciplinaires ? la systematique. pp. 293-303. Actes du 2e symposium international sur les Ombelliferes. Centre
!   Heracleum sphondylium Heracleum sphondylium L.   22+1S CromoCat 2015 Dempsey, R. E., Gornall, R. J. & Bailey, J. P. (1994). Contributions to a cytological catalogue of the British and Irish flora, 4. Watsonia 20: 63-66.
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Slovakia Database Májovský J. et al., 1974b: Index of chromosome numbers of Slovakian flora Part 4. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 23: 1-23.
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 British Isles cytology Maude, P.F. New Phytol. 38: 1-31 (1939)
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 British Isles cytology Dempsey, R.E., Gornall, R.J., Bailey, J.P. Watsonia 20: 63-66 (1994)
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 British Isles cytology Hollingsworth, P.M., Gornall, R.J., Bailey, J.P. Watsonia 19: 134-137 (1992)
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 British Isles cytology Weimarck, G., Stewart, F., Grace, J. Hereditas (Lund) 91: 117-127 (1979)
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 British Isles cytology Weimarck, G., Stewart, F., Grace, J. Hereditas (Lund) 91: 117-127 (1979)
  Heracleum sphondylium L. Heracleum sphondylium L. 11   Flora Iberica Nmeros cromosmicos para la flora espaola, 300-303. Pastor Daz, J.Lagascalia12(2): 279-280(1984).
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=3311
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=3313
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Flora Europaea Hakansson, 1953
  Heracleum sphondylium L. Heracleum sphondylium L.   22 Poland Database Skali?ska M., Ma?ecka J., Izmai?ow R. et al. 1974. Further studies in chromosome numbers of Polish Angiosperms. Tenth contribution. Acta Biol. Cracov. Ser. Bot. 17, 133-163.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 IPCN online Hollingsworth, P. M., R. J. Gornall & J. P. Bailey. 1992. Contribution to a cytological catalogue of the British and Irish flora, 2. Watsonia 19: 134–137.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 IPCN online Tonascia, N. 1992. Biosystematische Untersuchungen an Heracleum sphondylium s.l. in der Schweiz. Ber. Geobot. Inst. ETH Stiftung Rübel 58: 101–120.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 IPCN online Rostovtseva, T. S. 1979. Chromosome numbers of some species of the family Apiaceae Lindl. II. Bot. Zhurn. 64 (2): 227–232.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 IPCN online Strid, A. & R. Franzen. 1981. In Chromosome number reports LXXIII. Taxon 30: 829–842.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 IPCN online Skalinska, M. 1974. Further studies in chromosome numbers of Polish angiosperms. X. Acta Biol. Cracov., Ser. Bot. 17: 133–164.
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 IPCN73-74 MAJOVSKY, J., ET AL. 1974. Index of chromosome numbers of Slovakian Flora. (Part 4). Acta Fac. Rerum Nat. Univ. Comenianae Bot. 23: 1-23.
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Fedorov, 1974 Maude 1939, 1940
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Fedorov, 1974 Wulff 1939a
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Fedorov, 1974 P6lya 1950
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Fedorov, 1974 HAkansson 1953
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Fedorov, 1974 Quezel 1957
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Fedorov, 1974 Sorsa V. 1962
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 Darlington, 1955 Maude 1939
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 C-values database Siljak-Yakovlev S, Pustahija F, Šolic EM, Bogunic F, Muratovic E, Bašic N, Catrice O, Brown SC. 2010. Towards a genome size and chromosome number database of Balkan flora: C-values in 343 taxa with novel values for 242. Advanced Science Letters 3: 190-213
!   Heracleum sphondylium L. Heracleum sphondylium L. n = 11   Austria_checklist Wetschnig, W. & Leute, G. H. 1991. Chromosomenzahlen K?rntner Gef??pflanzen. Teil 2, Doldenbl?tler - Apiaceae = Umbelliferae. Linzer Biol.Beitr. 23[2], 457-481.°
!   Heracleum sphondylium L. Heracleum sphondylium L. n = 11   Austria_checklist Wetschnig, W. & Leute, G. H. 1991. Chromosomenzahlen K?rntner Gef??pflanzen. Teil 2, Doldenbl?tler - Apiaceae = Umbelliferae. Linzer Biol.Beitr. 23[2], 457-481.°
!   Heracleum sphondylium L. Heracleum sphondylium L.   2n = 24 Austria_checklist R. Mattick-Ehrensberger in Tischler, G. 1950. Die Chromosomenzahlen der Gef??pflanzen Mitteleuropas. p. 263. 's-Gravenhage, Uitgeverij Dr. W. Junk.
!   Heracleum sphondylium L. Heracleum sphondylium L.   2n = 22* Austria_checklist -- 1978. B chromosomes in Heracleum sphondylium s. lat. in Europe. Cauwet-Marc, A. M. and Carbonnier, J. Les Ombellif?res. Contributions pluridisciplinaires ? la systematique. pp. 293-303. Actes du 2e symposium international sur les Ombelliferes. Centre
!   Heracleum sphondylium L. Heracleum sphondylium L.   2n = 22 + 1 B* Austria_checklist -- 1978. B chromosomes in Heracleum sphondylium s. lat. in Europe. Cauwet-Marc, A. M. and Carbonnier, J. Les Ombellif?res. Contributions pluridisciplinaires ? la systematique. pp. 293-303. Actes du 2e symposium international sur les Ombelliferes. Centre
!   Heracleum sphondylium L. Heracleum sphondylium L.   2n = 22* Austria_checklist -- 1978. B chromosomes in Heracleum sphondylium s. lat. in Europe. Cauwet-Marc, A. M. and Carbonnier, J. Les Ombellif?res. Contributions pluridisciplinaires ? la systematique. pp. 293-303. Actes du 2e symposium international sur les Ombelliferes. Centre
  Heracleum sphondylium L. Heracleum sphondylium L.   c.22 Cave1963 Sorsa 1962
  Heracleum sphondylium L. Heracleum sphondylium L. 11   CromoCat 2015 Pastor Díaz, J. (1984). Números cromosómicos para la flora española, 300-303. Lagascalia 12(2): 279-280.
  Heracleum sphondylium L. Heracleum sphondylium L. 11   CromoCat 2015 Wetschnig, W. & G. H. Leute. (1991). Chromosomenzahlen Kärntner Gefässflanzen (Teil 2, Doldenblütler---Apiaceae = Umbelliferae). Linzer Biol. Beitr. 23: 457-481.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 CromoCat 2015 Strid, A. & R. Franzén. (1981). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports LXXIII]. Taxon 30(4): 829-842.
  Heracleum sphondylium L. Heracleum sphondylium L.   24 CromoCat 2015 Tischler, G. (1950). Die chromosomenzhalen der GefäBpflanzen Mitteleuropas. s'-Gravenhage: W. Junk. 263
!   Heracleum sphondylium L. Heracleum sphondylium L. 11   CromoCat 2015 Wetschnig, W. & G. H. Leute. (1991). Chromosomenzahlen Kärntner Gefässflanzen (Teil 2, Doldenblütler---Apiaceae = Umbelliferae). Linzer Biol. Beitr. 23: 457-481.
!   Heracleum sphondylium L. Heracleum sphondylium L.   11II+1BI CromoCat 2015 Weimarck, G. (1978). B chromosomes in Heracleum sphondylium s. lat. in Europe.-In: Cauwet-Marc A.M., Carbonnier J. (eds.): Les Ombellifères. Contributions pluridisciplinaires à la systematique. Actes du 2º symposium international sur les Ombelliferes. Per
!   Heracleum sphondylium L. Heracleum sphondylium L.   22* CromoCat 2015 Weimarck, G. (1978). B chromosomes in Heracleum sphondylium s. lat. in Europe.-In: Cauwet-Marc A.M., Carbonnier J. (eds.): Les Ombellifères. Contributions pluridisciplinaires à la systematique. Actes du 2º symposium international sur les Ombelliferes. Per
!   Heracleum sphondylium L. Heracleum sphondylium L.   22+1B* CromoCat 2015 Weimarck, G. (1978). B chromosomes in Heracleum sphondylium s. lat. in Europe.-In: Cauwet-Marc A.M., Carbonnier J. (eds.): Les Ombellifères. Contributions pluridisciplinaires à la systematique. Actes du 2º symposium international sur les Ombelliferes. Per
!   Heracleum sphondylium L. Heracleum sphondylium L.   22+2B* CromoCat 2015 Weimarck, G. (1978). B chromosomes in Heracleum sphondylium s. lat. in Europe.-In: Cauwet-Marc A.M., Carbonnier J. (eds.): Les Ombellifères. Contributions pluridisciplinaires à la systematique. Actes du 2º symposium international sur les Ombelliferes. Per
!   Heracleum sphondylium L. Heracleum sphondylium L.   22 CromoCat 2015 Lovka, M., Susnik, F., Löve, A. & Löve, D. (1972). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports XXXVI]. Taxon 21(2/3): 337-339.
  Heracleum sphondylium L. Heracleum sphondylium L.   22 IAPT/IOPB IAPT/IOPB Chromosome Data 27
!   Heracleum sphondylium None Heracleum sphondylium L.   22 CromoCat 2015 Löve, A. & Löve, D. (1982b). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports LXXV]. Taxon 31(2): 344-360.
!   Heracleum sphondylium None Heracleum sphondylium L.   22 CromoCat 2015 Siljak-Yakovlev, S., Pustahija, F., Šoli?, E.M., Boguni?, F., Muratovi?, E., Baši?, N., Catrice, O. & S. C. Brown. (2010). Towards a Genome Size and Chromosome Number Database of Balkan Flora: C-Values in 343 Taxa with Novel Values for 242 Advanced Scienc
!   Heracleum sphondylium subsp. alpinum (hartm.) ahlfv. Heracleum sphondylium subsp. alpinum (L.) Bonnier & Layens   22 Love, 1948 Maude 1939, 1940.
!   Heracleum sphondylium subsp. cyclocarpum P. H. Davis Heracleum cyclocarpum C. Koch   22+-4B Slovakia Database Murín A. & Májovský J., 1987: Karyological study of the Slovak flora XIX. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 34 [1986]: 3-20.
!   Heracleum sphondylium subsp. cyclocarpum P. H. Davis Heracleum cyclocarpum C. Koch   22+-4B Slovakia Database Murín A. & Májovský J., 1987: Karyological study of the Slovak flora XIX. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 34 [1986]: 3-20.
!   Heracleum sphondylium subsp. cyclocarpum P. H. Davis Heracleum cyclocarpum C. Koch   22 Slovakia Database Májovský J. et al., 1974b: Index of chromosome numbers of Slovakian flora Part 4. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 23: 1-23.
!   Heracleum sphondylium subsp. cyclocarpum P. H. Davis Heracleum cyclocarpum C. Koch   22@&4B IPCN online Murín, A. & J. Májovský. 1987. Karyological study of the Slovak flora XIX. Acta Fac. Rerum Nat. Univ. Comenianae, Bot. 34: 3–20.
!   Heracleum sphondylium subsp. cyclocarpum P. H. Davis Heracleum cyclocarpum C. Koch   22 IPCN online Murin, A. 1974. In Index of chromosome numbers of Slovakian flora. Part 4. Acta Fac. Rerum Nat. Univ. Comenianae, Bot. 23: 1–23.
!   Heracleum sphondylium subsp. cyclocarpum P. H. Davis Heracleum cyclocarpum C. Koch   22 IPCN73-74 MAJOVSKY, J., ET AL. 1974. Index of chromosome numbers of Slovakian Flora. (Part 4). Acta Fac. Rerum Nat. Univ. Comenianae Bot. 23: 1-23.
  Heracleum sphondylium subsp. lanatum (Michx.) Á. Löve & D. Löve Heracleum sphondylium var. lanatum (Michx.) Dorn   22 IPCN online Love, A. & D. Love. 1982. In: IOPB chromosome number reports LXXVI. Taxon 31: 583–587.
!   Heracleum sphondylium subsp. montanum (Schleich.) Briq. Heracleum sphondylium subsp. montanum (Schleich.) Briq.   22 IPCN72 LOVKA, M., F. SUSNIK, A. LOEVE, & D. LOEVE. 1972. In IOPB chromosome number reports XXXVI. Taxon 21: 333-346.
!   Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer 11   CHROBASE FAVARGER C. - 1973. Cytotaxonomie de quelques Orophytes des Abruzzes.. Acta Bot. Acad. Sci. Hung. , 19(1-4): 81-92.
!   Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer 11   IPCN73-74 FAVARGER, C. 1973. Cytotaxonomie de quelques orophytes des Abruzzes. Acta Bot. Acad. Sci. Hung. 18: 81-92.
!   Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 IPCN online Love, A. & D. Love. 1982. In: IOPB chromosome number reports LXXVI. Taxon 31: 583–587.
!   Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Leyens. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 CHROBASE LOEVE A., LOEVE D. - 1982. IOPB chromosome number reports LXXVI.. Taxon , 31(3): 583-587.
!   Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Leyens. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Leyens. Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. pyrenaicum None Heracleum sphondylium subsp. pyrenaicum (Lam.) Bonnier & Layens   22 CromoCat 2015 Löve, A. & Löve, D. (1982c). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports LXXVI]. Taxon 31(3): 583-587.
!   Heracleum sphondylium subsp. sibiricum (L.) Simonk. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 Flora Europaea Hakansson, 1953
  Heracleum sphondylium subsp. sibiricum (L.) Simonk. Heracleum sphondylium subsp. sibiricum (L.) Simonk.   22 IPCN online Arohonka, T. 1982. Chromosome counts of vascular plants of the island Seili in Nauvo, southwestern Finland. Turun Yliopiston Julkaisuja, Sar. A 2, Biol.-Geogr. 3: 1–12.
!   Heracleum sphondylium subsp. ternatum (Velen.) Brummitt Heracleum sphondylium subsp. ternatum (Velen.) Brummitt   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. ternatum (Velen.) Brummitt Heracleum sphondylium subsp. ternatum (Velen.) Brummitt   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. ternatum (Velen.) Brummitt Heracleum sphondylium subsp. ternatum (Velen.) Brummitt   22 CHROBASE WEIMARCK G. - 1978 (1977). B chromosomes in Heracleum sphondylium s. lat. in Europe.. Act. 2e Symp. Intern. Ombellifere (Perpignon) , : 293-303.
!   Heracleum sphondylium subsp. verticillatum (Pančić) Brummitt Heracleum sphondylium subsp. verticillatum (Pančić) Brummitt 11   IPCN75-78 Constance et al. 1976
  Heracleum ternatum Velen. Heracleum sphondylium subsp. ternatum (Velen.) Brummitt 11   IAPT/IOPB IAPT/IOPB Chromosome Data 10
  Heracleum ternatum Velen. Heracleum sphondylium subsp. ternatum (Velen.) Brummitt   22 IAPT/IOPB IAPT/IOPB Chromosome Data 27
!   Heracleum verticillatum Pančić Heracleum sphondylium subsp. verticillatum (Pančić) Brummitt   22 IPCN online Andreev, V. N. 1979. In IOPB chromosome number reports LXIV. Taxon 28: 397.